Vegetation Classes and Plant Communities (cont.)

Sagebrush Steppe.  Two sagebrush steppe classes are distinguished on the vegetation map:  Sagebrush Steppe on Lava and Sagebrush Steppe off Lava.  The distinction is largely a consequence of differences in soil reflectance rather than vegetation.  Soils overlying the basaltic uplands of the southern two-thirds of the INEEL tend to be darker than those of sagebrush-dominated communities off lava because of the presence of dark-colored pebbles or sand on the surface or because small patches of bare lava are exposed.  On the broken, irregular basalt topography, the soil surface often is sandy and friable and less likely to form the platy vesicular crusts that characterize many areas off the basalt flows.  This may influence  species composition.  For example, cheatgrass has invaded many of the communities on the basalt flows, but is rare or absent from some of the sagebrush dominated areas off the flows.  Soils in areas where cheatgrass is absent tend to be slightly saline and to form thick surface crusts.  Distinguishing between these two classes of sagebrush steppe also makes it easy to identify the extent of upland areas where shallow soils overlay relatively recent basalt flows.  One must keep in mind, however, that the classification is not completely accurate;  some sagebrush communities on the basalt flows may be classified as “off lava” and vice versa. 

Sagebrush steppe may be dominated by either Wyoming big sagebrush or basin big sagebrush, or by both.  The differential distributions of the two subspecies are related to gradients of soil texture (Shumar and Anderson 1986).  Basin big sagebrush occurs on sandy soils that are usually deep and well drained, whereas Wyoming big sagebrush tends to occur on fine-textured, shallow soils having limited depths of water infiltration.  Wyoming big sagebrush is the most abundant sagebrush at the INEEL, but extensive patches of basin big sagebrush occur on the eastern basalt uplands (Shumar 1983).  Pockets of basin big sagebrush within areas generally dominated by Wyoming big sagebrush occur on the lee sides of lava ridges where sand accumulates, within former channels of the Big Lost River, and on the linear sand dunes near the Mud Lake basin.  The influence of these linear dunes is readily apparent on the vegetation map.

Some areas of sagebrush steppe are dominated by one of the species of “low sage,” black sagebrush (Artemisia nova) or Artemisia arbuscula.  Black sagebrush typically occurs on shallow, stony, calcareous or limestone-derived soils (Shultz 1983, Hironaka 1979), although it also occurs on dry, windswept ridges and on shallow soils overlying basalt.  A. arbuscula also occurs on shallow soils, but it is more of an upland species, often found on foothill slopes.

Aside from the dominance of sagebrush, the sagebrush steppe communities do not have a unique species composition.  However, most sagebrush steppe has an abundance of perennial grasses, and the combined cover of perennial grasses can approach that of the sagebrush.  These are the same grasses that occur in the grasslands described earlier.  In fact, the only apparent difference between many areas classified as sagebrush steppe and others classified as grasslands is the density of sagebrush.  In some areas, this reflects fire history.  Sagebrush is killed by fire and must recolonize burned sites from seed.  Most perennial grasses and forbs resprout following fire from roots or other organs that are protected below ground.  Therefore, these herbaceous species can become the dominants following fire and it may be decades before sagebrush again reaches a high density.  Such areas likely will be classified as grasslands.

Shrubs other than sagebrush occur in most of these steppes.  Green rabbitbrush is essentially ubiquitous;  other common shrubs include winterfat, prickly phlox, and spiny hopsage.

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